精灵动画实现中的background-position像素是如何计算的？

@-webkit-keyframes person-slow {
        0% {
            background-position: -0px -291px;
        }
        25% {
            background-position: -602px -0px;
        }
        50% {
            background-position: -302px -291px;
        }
        75% {
            background-position: -151px -291px;
        }
        100% {
            background-position: -0px -291px;
        }
    }